The 1st one (no cap) requires a header tank to fill / top up the system. The 2nd one (cap and hose) has a built in cap! and you can fill up your coolant system from that. The hose is an overflow in case the system is over filled / pressured. That might benefit an expansion bottle, but not sure if its an absolute requirement (it wasn't on my old MGB)
Edit: an expansion tank isn't the same as a header tank. The expansion tank takes any overflow from a system and isn't pressurised. A header tank is part of the system and is pressurised.
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